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<front>
<journal-meta>
<journal-id journal-id-type="publisher">FR</journal-id>
<journal-title-group>
<journal-title>Fossil Record</journal-title>
<abbrev-journal-title abbrev-type="publisher">FR</abbrev-journal-title>
<abbrev-journal-title abbrev-type="nlm-ta">Foss. Rec.</abbrev-journal-title>
</journal-title-group>
<issn pub-type="epub">2193-0074</issn>
<publisher><publisher-name>Copernicus Publications</publisher-name>
<publisher-loc>Göttingen, Germany</publisher-loc>
</publisher>
</journal-meta>
<article-meta>
<article-id pub-id-type="doi">10.5194/fr-17-33-2014</article-id>
<title-group>
<article-title>Osteoderm distribution has low impact  on the centre of mass of stegosaurs</article-title>
</title-group>
<contrib-group><contrib contrib-type="author" xlink:type="simple"><name name-style="western"><surname>Mallison</surname>
<given-names>H.</given-names>
</name>
<xref ref-type="aff" rid="aff1">
<sup>1</sup>
</xref>
</contrib>
</contrib-group><aff id="aff1">
<label>1</label>
<addr-line>Museum für Naturkunde – Leibniz-Institut für Evolutions- und Biodiversitätsforschung an der Humboldt-Universität zu Berlin, Berlin, Germany</addr-line>
</aff>
<pub-date pub-type="epub">
<day>07</day>
<month>03</month>
<year>2014</year>
</pub-date>
<volume>17</volume>
<issue>1</issue>
<fpage>33</fpage>
<lpage>39</lpage>
<permissions>
<copyright-statement>Copyright: &#x000a9; 2014 H. Mallison</copyright-statement>
<copyright-year>2014</copyright-year>
<license license-type="open-access">
<license-p>This work is licensed under the Creative Commons Attribution 3.0 Unported License. To view a copy of this licence, visit <ext-link ext-link-type="uri"  xlink:href="https://creativecommons.org/licenses/by/3.0/">https://creativecommons.org/licenses/by/3.0/</ext-link></license-p>
</license>
</permissions>
<self-uri xlink:href="https://fr.copernicus.org/articles/17/33/2014/fr-17-33-2014.html">This article is available from https://fr.copernicus.org/articles/17/33/2014/fr-17-33-2014.html</self-uri>
<self-uri xlink:href="https://fr.copernicus.org/articles/17/33/2014/fr-17-33-2014.pdf">The full text article is available as a PDF file from https://fr.copernicus.org/articles/17/33/2014/fr-17-33-2014.pdf</self-uri>
<abstract>
<p>It has been hypothesized that the pronounced differences of stegosaur
humeral shapes, with large forms having more slender and small forms having
more robust humeri, may be explained by a difference in relative centre of
mass (COM) placement caused by differing distributions of osteoderms. To
test this hypothesis, digital 3-D models of the bones and osteoderms of the
Tanzanian stegosaur &lt;i&gt;Kentrosaurus aethiopicus&lt;/i&gt; and of the North
American stegosaur &lt;i&gt;Stegosaurus armatus&lt;/i&gt; were used to create
a 3-D computer-aided design life reconstruction. On these models osteoderm
placement was varied drastically, recreating both existing and hypothetical
forms. These models show that COM position varies somewhat with realistic
osteoderm distributions, but insufficiently to explain major differences in
humeral shape. The uniform weight distribution between forelimbs and hindlimbs 
found between the two taxa also casts doubt on the hypothesis that
differences in relative COM position caused by other factors than osteoderm
distribution can explain differences in humeral robustness.</p>
</abstract>
<counts><page-count count="7"/></counts>
</article-meta>
</front>
<body/>
<back>
<ref-list>
<title>References</title>
<ref id="ref1">
<label>1</label><mixed-citation publication-type="other" xlink:type="simple">Alexander, R. M.: Dynamics of dinosaurs and other extinct giants, Columbia University Press, New York, 1989.</mixed-citation>
</ref>
<ref id="ref2">
<label>2</label><mixed-citation publication-type="other" xlink:type="simple">Allen, V. R., Paxton, H. and Hutchinson, J. R.: Variation of center of mass estimates for extant sauropsids and its importance for reconstructing inertial properties of extinct archosaurs, Anat. Rec., 292, 1442–1461, 2009.</mixed-citation>
</ref>
<ref id="ref3">
<label>3</label><mixed-citation publication-type="other" xlink:type="simple">Bakker, R. T.: The Dinosaur Heresis, William Morrow and Co., New York, 1986.</mixed-citation>
</ref>
<ref id="ref4">
<label>4</label><mixed-citation publication-type="other" xlink:type="simple">Butler, R. J. and Barrett, P. M.: Reassessment of the evidence for postcranial skeletal pneumaticity in Triassic archosaurs, and the early evolution of the avian respiratory system, PLOS ONE, 7, e34094, &lt;a href=&quot;http://dx.doi.org/10.1371/journal.pone.0034094&quot;&gt;https://doi.org/10.1371/journal.pone.0034094&lt;/a&gt;, 2012.</mixed-citation>
</ref>
<ref id="ref5">
<label>5</label><mixed-citation publication-type="other" xlink:type="simple">Carpenter, K.: Armor of &lt;i&gt;Stegosaurus stenops&lt;/i&gt;, and the taphonomic history of a new specimen from Garden Park Colorado, Up. Jurass. Morrison Form. Interdiscip. Study Part 1 Mod. Geol., 22, 127–144, 1998.</mixed-citation>
</ref>
<ref id="ref6">
<label>6</label><mixed-citation publication-type="other" xlink:type="simple">Colbert, E. H.: The weights of dinosaurs, Am. Mus. Novit., 2076, 1–17, 1962.</mixed-citation>
</ref>
<ref id="ref7">
<label>7</label><mixed-citation publication-type="other" xlink:type="simple">Currey, J. D.: Bones: Structure and Mechanics, Princeton University Press, New Jersey, 2002.</mixed-citation>
</ref>
<ref id="ref8">
<label>8</label><mixed-citation publication-type="other" xlink:type="simple">Falkingham, P. L.: Acquisition of high resolution three-dimensional models using free, open-source, photogrammetric software, Palaeontol. Electron., 15.1.1T, 2011.</mixed-citation>
</ref>
<ref id="ref9">
<label>9</label><mixed-citation publication-type="other" xlink:type="simple">Galton, P. M.: The postcranial anatomy of stegosaurian dinosaur &lt;i&gt;Kentrosaurus&lt;/i&gt; from the Upper Jurassic of Tanzania, East Africa, Geol. Palaeontol., 15, 139–165, 1982.</mixed-citation>
</ref>
<ref id="ref10">
<label>10</label><mixed-citation publication-type="other" xlink:type="simple">Galton, P. M. and Upchurch, P.: Stegosauria, in: The Dinosauria, 2nd Edition, edited by: Weishampel, D. B., Dodson, P., and Osmolska, H., University of California Press, Berkeley, 343–362, 2004.</mixed-citation>
</ref>
<ref id="ref11">
<label>11</label><mixed-citation publication-type="other" xlink:type="simple">Gilmore, C. W.: Osteology of the armored Dinosauria in the United States National Museum, with special reference to the genus &lt;i&gt;Stegosaurus&lt;/i&gt;, U.S. Natl. Mus. Bull., 89, 1–136, 1914.</mixed-citation>
</ref>
<ref id="ref12">
<label>12</label><mixed-citation publication-type="other" xlink:type="simple">Gunga, H.-C., Suthau, T., Bellmann, A., Friedrich, A., Schwanebeck, T., Stoinski, S., Trippel, T., Kirsch, K. and Hellwich, O.: Body mass estimations for &lt;i&gt;Plateosaurus engelhardti&lt;/i&gt; using laser scanning and 3D reconstruction methods, Naturwissenschaften, 94, 623–630, 2007.</mixed-citation>
</ref>
<ref id="ref13">
<label>13</label><mixed-citation publication-type="other" xlink:type="simple">Henderson, D. M.: Estimating the masses and centers of mass of extinct animals by 3D mathematical slicing, Paleobiology, 25, 88–106, 1999.</mixed-citation>
</ref>
<ref id="ref14">
<label>14</label><mixed-citation publication-type="other" xlink:type="simple">Hennig, E.: &lt;i&gt;Kentrosaurus aethiopicus&lt;/i&gt; der Stegosauride des Tendaguru, Sitzungsberichte Ges. Naturforschender Freunde Zu Berl., 1915, 219–247, 1915.</mixed-citation>
</ref>
<ref id="ref15">
<label>15</label><mixed-citation publication-type="other" xlink:type="simple">Hennig, E.: &lt;i&gt;Kentrurosaurus aethiopicus&lt;/i&gt; Die Stegosaurier-Funde vom Tendaguru, Deutsch-Ostafrika, Palaeontographica, Supplement 7, 101–254, 1925.</mixed-citation>
</ref>
<ref id="ref16">
<label>16</label><mixed-citation publication-type="other" xlink:type="simple">Janensch, W.: Ein aufgestelltes Skelett des Stegosauriers &lt;i&gt;Kentrurosaurus aethiopicus&lt;/i&gt; HENNIG 1915 aus den Tendaguru-Schichten Deutsch-Ostafrikas, Palaeontographica, Supplement 7, 255–276, 1925.</mixed-citation>
</ref>
<ref id="ref17">
<label>17</label><mixed-citation publication-type="other" xlink:type="simple">Maidment, S. C. R., Norman, D. B., Barret, P. M., and Upchurch, P.: Systematics and phylogeny of Stegosauria (Dinosauria: Ornithischia), J. Syst. Palaeontol., 6, 367–407, &lt;a href=&quot;http://dx.doi.org/10.1017/S1477201908002459&quot;&gt;https://doi.org/10.1017/S1477201908002459&lt;/a&gt;, 2008.</mixed-citation>
</ref>
<ref id="ref18">
<label>18</label><mixed-citation publication-type="other" xlink:type="simple">Maidment, S. C. R., Linton, D. H., Upchurch, P., and Barrett, P. M.: Limb-bone scaling indicates diverse stance and gait in quadrupedal ornithischian dinosaurs, PLOS ONE, 7, e36904, &lt;a href=&quot;http://dx.doi.org/10.1371/journal.pone.0036904&quot;&gt;https://doi.org/10.1371/journal.pone.0036904&lt;/a&gt;, 2012.</mixed-citation>
</ref>
<ref id="ref19">
<label>19</label><mixed-citation publication-type="other" xlink:type="simple">Maidment, S. C. R., Bates, K. T., and Barrett, P. M.: Three-dimensional computational modeling of locomotor muscle moment arms in &lt;i&gt;Edmontosaurus&lt;/i&gt; (Dinosauria: Hadrosauridae) and comparisons with other archosaurs, in: Hadrosaurs, edited by: Evans, D. E. and Eberth, D. C., Indiana University Press, in press, 2014.</mixed-citation>
</ref>
<ref id="ref20">
<label>20</label><mixed-citation publication-type="other" xlink:type="simple">Maidment, S. C. R., Bates, K. T., Falkingham, P. L., VanBuren, C., Arbour, V. M., and Barrett, P. M.: Locomotion in ornithischian dinosaurs: an assessment using three-dimensional computational modeling, Biol. Rev., &lt;a href=&quot;http://dx.doi.org/10.1111/brv.12071&quot;&gt;https://doi.org/10.1111/brv.12071&lt;/a&gt;, 2013b.</mixed-citation>
</ref>
<ref id="ref21">
<label>21</label><mixed-citation publication-type="other" xlink:type="simple">Mallison, H.: CAD assessment of the posture and range of motion of &lt;i&gt;Kentrosaurus aethiopicus&lt;/i&gt; Hennig 1915, Swiss J. Geosci., 103, 211–233, &lt;a href=&quot;http://dx.doi.org/10.1007/s00015-010-0024-2&quot;&gt;https://doi.org/10.1007/s00015-010-0024-2&lt;/a&gt;, 2010.</mixed-citation>
</ref>
<ref id="ref22">
<label>22</label><mixed-citation publication-type="other" xlink:type="simple">Mallison, H.: Defense capabilities of &lt;i&gt;Kentrosaurus aethiopicus&lt;/i&gt; Hennig, 1915, Palaeontol. Electron., 14, 10A, 2011a.</mixed-citation>
</ref>
<ref id="ref23">
<label>23</label><mixed-citation publication-type="other" xlink:type="simple">Mallison, H.: Rearing Giants – kinetic/dynamic modeling of sauropod bipedal and tripodal poses, in: Biology of the Sauropod Dinosaurs: Understanding the life of giants, edited by: Klein, N., Remes, K., Gee, C. T., and Sander, P. M., Indiana University Press, Bloomington, 237–250, 2011b.</mixed-citation>
</ref>
<ref id="ref24">
<label>24</label><mixed-citation publication-type="other" xlink:type="simple">Mallison, H.: The real lectotype of &lt;i&gt;Kentrosaurus aethiopicus&lt;/i&gt; Hennig 1915, Neues Jahrb. für Geol. Paläontol. Abh., 259, 197–206, 2011c.</mixed-citation>
</ref>
<ref id="ref25">
<label>25</label><mixed-citation publication-type="other" xlink:type="simple">Mateus, O., Maidment, S. C. R., and Christiansen, N. A.: A new long-necked &quot;sauropod-mimic&quot; stegosaur and the evolution of the plated dinosaurs, Proc. R. Soc. B Biol. Sci., 276, 1815–1821, &lt;a href=&quot;http://dx.doi.org/10.1098/rspb.2008.1909&quot;&gt;https://doi.org/10.1098/rspb.2008.1909&lt;/a&gt;, 2009.</mixed-citation>
</ref>
<ref id="ref26">
<label>26</label><mixed-citation publication-type="other" xlink:type="simple">Ouyang, H.: [Discovery of &lt;i&gt;Gigantspinosaurus sichuanensis&lt;/i&gt; and its scapular spine orientation.], Abstr. Summ. Youth Acad. Symp. New Discov. Ideas Stratigr. Paleontol. Nanjing Dec 1992, 47–49, 1992.</mixed-citation>
</ref>
<ref id="ref27">
<label>27</label><mixed-citation publication-type="other" xlink:type="simple">Siber, H. J. and Möckli, U.: The Stegosaurs of the Sauriermuseum Aathal, Sauriermuseum Aathal, Aathal, Switzerland, 2009.</mixed-citation>
</ref>
</ref-list>
</back>
</article>