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<front>
<journal-meta>
<journal-id journal-id-type="publisher">FR</journal-id>
<journal-title-group>
<journal-title>Fossil Record</journal-title>
<abbrev-journal-title abbrev-type="publisher">FR</abbrev-journal-title>
<abbrev-journal-title abbrev-type="nlm-ta">Foss. Rec.</abbrev-journal-title>
</journal-title-group>
<issn pub-type="epub">2193-0074</issn>
<publisher><publisher-name>Wiley</publisher-name>
<publisher-loc>Göttingen, Germany</publisher-loc>
</publisher>
</journal-meta>
<article-meta>
<article-id pub-id-type="doi">10.1002/mmng.20000030109</article-id>
<title-group>
<article-title>On the systematic and stratigraphic significance of pterosaurs from the Lower Cretaceous Yixian Formation (Jehol Group) of Liaoning, China</article-title>
</title-group>
<contrib-group><contrib contrib-type="author" xlink:type="simple"><name name-style="western"><surname>Unwin</surname>
<given-names>D. M.</given-names>
</name>
<xref ref-type="aff" rid="aff1">
<sup>1</sup>
</xref>
</contrib>
<contrib contrib-type="author" xlink:type="simple"><name name-style="western"><surname>Lü</surname>
<given-names>J.</given-names>
</name>
<xref ref-type="aff" rid="aff2">
<sup>2</sup>
</xref>
</contrib>
<contrib contrib-type="author" xlink:type="simple"><name name-style="western"><surname>Bakhurina</surname>
<given-names>N. N.</given-names>
</name>
<xref ref-type="aff" rid="aff3">
<sup>3</sup>
</xref>
</contrib>
</contrib-group><aff id="aff1">
<label>1</label>
<addr-line>Museum für Naturkunde, Zentralinstitut der Humboldt-Universität zu Berlin, Invalidenstraße 43, 10115 Berlin, Germany</addr-line>
</aff>
<aff id="aff2">
<label>2</label>
<addr-line>Institute of Vertebrate Palaeontology and Palaeoanthropology, Academica Sinica, 142 Xi-Zhi-Men-Wai Street, P.O. Box 643, Beijing 100044, China</addr-line>
</aff>
<aff id="aff3">
<label>3</label>
<addr-line>Department of Earth Sciences, University of Bristol, Queen&apos;s Road, Bristol BS8 1RJ, UK</addr-line>
</aff>
<pub-date pub-type="epub">
<day>01</day>
<month>01</month>
<year>2000</year>
</pub-date>
<volume>3</volume>
<issue>1</issue>
<fpage>181</fpage>
<lpage>206</lpage>
<permissions>
<copyright-statement>Copyright: &#x000a9; 2000 D. M. Unwin et al.</copyright-statement>
<copyright-year>2000</copyright-year>
<license license-type="open-access">
<license-p>This work is licensed under the Creative Commons Attribution 3.0 Unported License. To view a copy of this licence, visit <ext-link ext-link-type="uri"  xlink:href="https://creativecommons.org/licenses/by/3.0/">https://creativecommons.org/licenses/by/3.0/</ext-link></license-p>
</license>
</permissions>
<self-uri xlink:href="https://fr.copernicus.org/articles/3/181/2000/fr-3-181-2000.html">This article is available from https://fr.copernicus.org/articles/3/181/2000/fr-3-181-2000.html</self-uri>
<self-uri xlink:href="https://fr.copernicus.org/articles/3/181/2000/fr-3-181-2000.pdf">The full text article is available as a PDF file from https://fr.copernicus.org/articles/3/181/2000/fr-3-181-2000.pdf</self-uri>
<abstract>
<p>A reassessment of the systematic relationships of pterosaurs from the Lower Cretaceous Yixian Formation of Liaoning Province, China, shows that &lt;i&gt;Dendrorhynchoides&lt;/i&gt; should be reassigned to the Anurognathidae (“Rhamphorhynchoidea”) and that &lt;i&gt;Eosipterus&lt;/i&gt; possibly belongs within Ctenochasmatidae (Pterodactyloidea). These pterosaurs formed an integral part of a diverse community that inhabited lowland terrestrial environments in the region of northeast China in the Early Cretaceous. A new compilation of data for the Lower Cretaceous hints at a broad differentiation between pterosaurs that lived in continental habitats (anurognathids, ctenochasmatoids, dsungaripteroids) and those that frequented marine environments (ornithocheiroids). Moreover, there is evidence of further differentiation within continental habitats, between pterosaurs living in lowland and coastal regions (anurognathids. ctenochasmatoids) and those living in more inland environments (dsungaripteroids). The temporal and geographical range extensions for high rank taxa that are implied by the Yixian pterosaurs further emphasise the incompleteness and unevenness of the pterosaur fossil record and its unreliability for biostratigraphic zonation.
&lt;br&gt;&lt;br&gt;
Eine Neubewertung der systematischen Stellung der Flugsaurier von der unterkretazischen Yixian-Formation der Provinz Liaoning, China, zeigt, dass &lt;i&gt;Dendrorhynchoides&lt;/i&gt; den Anurognathiden (“Rhamphorhynchoidea”) zugeordnet werden kann und dass &lt;i&gt;Eosipterus&lt;/i&gt; vermutlich zu den Ctenochasmatiden (Pterodactyloidea) gehört. Diese beiden Flugsaurier bilden einen integralen Bestandteil einer diversen Fauna, die in der Unteren Kreide ein terrestrisches Flachland-Ökosystem im Nordosten Chinas bewohnte. Fasst man die für die Untere Kreide verfügbaren Daten zusammen, so zeigt sich eine weitgehende Differenzierung zwischen Flugsauriern, die überwiegend in kontinentalen Ökosystemen lebten (Anurognathidae, Ctenochasmatoidea, Dsungaripteroidea) und jenen, die auch oft in marinen Bereichen auftreten (Ornithocheiroidea). Darüber hinaus gibt es auch Hinweise auf eine Differenzierung innerhalb der kontinentalen Habitate, zwischen Pterosauriern, die sich in den Ebenen und küstennahen Bereichen aufhielten (Anurognathidae, Ctenochasmatoidea) und den Bewohnern von mehr küstenfernen Ökosystemen (Dsungaripteroidea). Die von den Taxa der Yixian-Formation angezeigte Erweiterung der stratigraphischen und geographischen Reichweite für Taxa höheren Ranges unterstreichen die Unvollständigkeit und Unausgewogenheit des Fossilberichtes der Flugsaurier und seine Unzulänglichkeit für biostratigraphische Zonierungen.
&lt;br&gt;&lt;br&gt;
doi:&lt;a href=&quot;http://dx.doi.org/10.1002/mmng.20000030109&quot; target=&quot;_blank&quot;&gt;10.1002/mmng.20000030109&lt;/a&gt;</p>
</abstract>
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