Fossil Record
Fossil Record
Fossil Record
Articles | Volume 24, issue 1
Articles | Volume 24, issue 1
https://doi.org/10.5194/fr-24-101-2021
© Author(s) 2021. This work is distributed under
the Creative Commons Attribution 4.0 License.
https://doi.org/10.5194/fr-24-101-2021
© Author(s) 2021. This work is distributed under
the Creative Commons Attribution 4.0 License.
Articles | Volume 24, issue 1
Research article
 | 
23 Apr 2021
Research article |  | 23 Apr 2021

Extensive vertebrate tracksite from the Upper Red Formation (middle–late Miocene), west Zanjan, northwestern Iran

Nasrollah Abbassi, Masoumeh Khoshyar, Spencer G. Lucas, and Farid Esmaeili

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Latest update: 21 Jul 2026
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Short summary
The Chehrabad area, NW Iran, is known as the historical site of Saltmen mine dated to the Achaemenid and Sassanid eras. We, however, discovered animal footprints in NW Chehrabad (ca. 3 km NW of the Saltmen mine) on a very large slab. These footprints belong to the Miocene age and include canid, felid and bird tracks as old as 10.7 Ma. The canid footprint record establishes the late Miocene presence of canids on the Iranian Plateau as part of the Eurasia-wide “Eucyon event”.
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