Articles | Volume 24, issue 1
https://doi.org/10.5194/fr-24-193-2021
© Author(s) 2021. This work is distributed under
the Creative Commons Attribution 4.0 License.
the Creative Commons Attribution 4.0 License.
https://doi.org/10.5194/fr-24-193-2021
© Author(s) 2021. This work is distributed under
the Creative Commons Attribution 4.0 License.
the Creative Commons Attribution 4.0 License.
Sheltered preservation in Ordovician trilobites
Oldřich Fatka
CORRESPONDING AUTHOR
Institute of Geology and Palaeontology, Faculty of Science, Charles
University, Albertov 6, Prague 2, 128 43, Czech Republic
Petr Budil
Czech Geological Survey, Klárov 3, Prague 1, 118 21, Czech
Republic
Petr Kraft
Institute of Geology and Palaeontology, Faculty of Science, Charles
University, Albertov 6, Prague 2, 128 43, Czech Republic
Related subject area
Evolution and phylogeny
The life cycle in late Paleozoic eryopid temnospondyls: developmental variation, plasticity and phylogeny
A unique, large-sized stem Odonata (Insecta) found in the early Pennsylvanian of New Brunswick (Canada)
Determining the gait of Miocene, Pliocene, and Pleistocene horses from fossilized trackways
†Cretolixon – a remarkable new genus of rhopalosomatid wasps (Hymenoptera: Vespoidea: Rhopalosomatidae) from chemically tested, mid-Cretaceous Burmese (Kachin) amber supports the monophyly of Rhopalosomatinae
The youngest occurrence of embolomeres (Tetrapoda: Anthracosauria) from the Sunjiagou Formation (Lopingian, Permian) of North China
Sampling fossil floras for the study of insect herbivory: how many leaves is enough?
A new species of baenid turtle from the Early Cretaceous Lakota Formation of South Dakota
Quantitative analysis of suture lines in Carboniferous ammonoids
Osseous anomalies of the cryptobranchid Eoscapherpeton asiaticum (Amphibia: Caudata) from the Late Cretaceous of Uzbekistan
Growth patterns, sexual dimorphism, and maturation modeled in Pachypleurosauria from Middle Triassic of central Europe (Diapsida: Sauropterygia)
New information on the Cenomanian bothremydid turtle Algorachelus based on new, well-preserved material from Spain
A case study of developmental palaeontology in Stereosternum tumidum (Mesosauridae, Parareptilia)
Postcrania of large dissorophid temnospondyls from Richards Spur, Oklahoma
Description of the skeleton of the fossil beaked whale Messapicetus gregarius: searching potential proxies for deep-diving abilities
Foramina in plesiosaur cervical centra indicate a specialized vascular system
The wasp larva's last supper: 100 million years of evolutionary stasis in the larval development of rhopalosomatid wasps (Hymenoptera: Rhopalosomatidae)
The fossil history of pseudoscorpions (Arachnida: Pseudoscorpiones)
The blueprint of the Amphiesmenoptera – Tarachoptera, a new order of insects from Burmese amber (Insecta, Amphiesmenoptera)
Modeling the physiology of the aquatic temnospondyl Archegosaurus decheni from the early Permian of Germany
Turtle shell bone and osteoderm histology of Mesozoic and Cenozoic stem-trionychian Adocidae and Nanhsiungchelyidae (Cryptodira: Adocusia) from Central Asia, Mongolia, and North America
Reinvestigation of Protelytron permianum (Insecta; Early Permian; USA) as an example for applying reflectance transformation imaging to insect imprint fossils
A new species of Cyclotosaurus (Stereospondyli, Capitosauria) from the Late Triassic of Bielefeld, NW Germany, and the intrarelationships of the genus
New remarkable Late Jurassic teleosts from southern Germany: Ascalaboidae n. fam., its content, morphology, and phylogenetic relationships
New specimen of Cacops woehri indicates differences in the ontogenetic trajectories among cacopine dissorophids
The stratigraphic importance of the brontothere (cf. Diplacodon elatus) in the Brennan Basin Member of the Duchesne River Formation of Utah
Do tracks yield reliable information on gaits? – Part 1: The case of horses
Rainer R. Schoch
Foss. Rec., 24, 295–319, https://doi.org/10.5194/fr-24-295-2021, https://doi.org/10.5194/fr-24-295-2021, 2021
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Eryopids were 2 m large relatives of modern amphibians that formed top predators in the Paleozoic. They are found in aquatic deposits and apparently preyed on fishes. Their life cycle is preserved in a single species, Onchiodon labyrinthicus, from eastern Germany. It existed in a stressed ecosystem, and the broad variation in the studied sample correlates with seasonal fluctuations. Eryopids evolved into increasingly large, heavily built inhabitants of streams and seasonal water bodies.
Olivier Béthoux, Rowan E. Norrad, Matthew R. Stimson, Olivia A. King, Luke F. Allen, Isabelle Deregnaucourt, Steven J. Hinds, Jake H. Lewis, and Jörg W. Schneider
Foss. Rec., 24, 207–221, https://doi.org/10.5194/fr-24-207-2021, https://doi.org/10.5194/fr-24-207-2021, 2021
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A specimen of an extinct relative of damsel- and dragonflies was discovered from ca. 310-million-years-old strata from New Brunswick (Canada). It is composed of an incomplete hindwing belonging to a new species whose wingspan was about 35 cm. Despite its incompleteness, it composes a useful addition to our knowledge of the early evolution of the group, owing to its unique combination of traits.
Alan Vincelette
Foss. Rec., 24, 151–169, https://doi.org/10.5194/fr-24-151-2021, https://doi.org/10.5194/fr-24-151-2021, 2021
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This study compares footprints made by modern horses with two fossil trackways. One shows a Pleistocene horse in a gallop of 9 m/s, perhaps fleeing predators or migrating. More intriguing is the trackway of a Miocene horse with diagonal couplets in a gait of 2 m/s. This best matches the artificial gait of the rack/tölt only found in breeds such as the Saddlebred, Icelandic, and Paso, providing more evidence that early horses possessed a greater variety of useful gaits than most modern horses.
Volker Lohrmann, Qi Zhang, Peter Michalik, Jeremy Blaschke, Patrick Müller, Laurent Jeanneau, and Vincent Perrichot
Foss. Rec., 23, 215–236, https://doi.org/10.5194/fr-23-215-2020, https://doi.org/10.5194/fr-23-215-2020, 2020
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A new mid-Cretaceous rhopalosomatid wasp, Cretolixon alatum Lohrmann gen. et sp. nov., is described from Burmese (Kachin) amber. The new genus has a unique mixture of characters, some of which are only known from the recent brachypterous genus Olixon and others of which are known only from the recent macropterous genera. Thus, Cretolixon not only provides further evidence for the monophyly of the family but also contributes evidence for the monophyly of the Rhopalosomatinae.
Jianye Chen and Jun Liu
Foss. Rec., 23, 205–213, https://doi.org/10.5194/fr-23-205-2020, https://doi.org/10.5194/fr-23-205-2020, 2020
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In this paper, we describe a new species of embolomere from 255-million-year-old fossil beds from northern China. This group of ancient animals is closely related to amniotes (reptiles, birds, and mammals) and was previously thought to have gone extinct around 273 million years ago. The new discovery indicates that North China, with its ancient tropical rain forest, became a last reservoir for these animals right before the end-Permian mass extinction.
Sandra R. Schachat, S. Augusta Maccracken, and Conrad C. Labandeira
Foss. Rec., 23, 15–32, https://doi.org/10.5194/fr-23-15-2020, https://doi.org/10.5194/fr-23-15-2020, 2020
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Insects have been feeding on plants for over 300 million years. The fossil record preserves evidence of this behavior: it is possible to examine a fossilized leaf and see the spots where insects ate it or laid their eggs in it. The study of these insect
traceson leaves shows us how insects have adapted to changing climates. Here, we address the question of how many leaves from a single fossil locality must be examined to permit statistically robust comparisons in deep time.
Walter G. Joyce, Yann Rollot, and Richard L. Cifelli
Foss. Rec., 23, 1–13, https://doi.org/10.5194/fr-23-1-2020, https://doi.org/10.5194/fr-23-1-2020, 2020
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Baenids are a group of fossil turtles with a rich fossil record that spans from the Late Cretaceous (Campanian) to Eocene of North America. Only a few fossils are known that document the early evolution of the group. We here describe a new species of baenid, Lakotemys australodakotensis, from the Early Cretaceous (Berriasian to Valanginian) Lakota Formation of South Dakota, USA. Phylogenetic analysis suggests that this is the oldest known baenid turtle.
Michael Wegerer, Kenneth De Baets, and Dieter Korn
Foss. Rec., 21, 223–236, https://doi.org/10.5194/fr-21-223-2018, https://doi.org/10.5194/fr-21-223-2018, 2018
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Two morphometric methods are applied for the analysis of suture lines in Early Carboniferous ammonoids: (1) classic metric data using multivariate statistic methods and (2) outline data are analysed using the elliptic Fourier analysis. Both methods lead to similar results and demonstrate ontogenetic and phylogenetic trends in these ammonoids: (1) a general decrease in the amplitude of lobes and saddles, (2) a proportional widening of the external lobe, and (3) a heightening of the median saddle.
Pavel Skutschas, Veniamin Kolchanov, Elizaveta Boitsova, and Ivan Kuzmin
Foss. Rec., 21, 159–169, https://doi.org/10.5194/fr-21-159-2018, https://doi.org/10.5194/fr-21-159-2018, 2018
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Paleopathologies among fossil members of modern groups of amphibians are poorly documented. We describe pathologies of the fossil salamander Eoscapherpeton asiaticum from the Late Cretaceous of Uzbekistan. These pathologies can result from trauma, infection from trauma and congenital disorders. The occurrence of several traumatic femoral pathologies in Eoscapherpeton could result from intraspecific aggressive behavior. Bone pathologies are described for the first time in fossil salamanders.
Nicole Klein and Eva Maria Griebeler
Foss. Rec., 21, 137–157, https://doi.org/10.5194/fr-21-137-2018, https://doi.org/10.5194/fr-21-137-2018, 2018
Adán Pérez-García
Foss. Rec., 21, 119–135, https://doi.org/10.5194/fr-21-119-2018, https://doi.org/10.5194/fr-21-119-2018, 2018
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The Spanish Algorachelus peregrinus is the oldest pleurodiran turtle known in the Northern Hemisphere. New material is presented, including several shells. The patterns by which some of them were disarticulated are analyzed. Several pathologies are identified. The knowledge about the anatomy and intraspecific variability is improved. Algorachelus is recognized in the Middle East and North America. The oldest known dispersal event of Pleurodira from Gondwana to Laurasia is characterized.
Constanze Bickelmann and Linda A. Tsuji
Foss. Rec., 21, 109–118, https://doi.org/10.5194/fr-21-109-2018, https://doi.org/10.5194/fr-21-109-2018, 2018
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Mesosaurs were reptiles that lived around 280 mya. They were the first land reptile group to return to the water. Study of a fossil from the Museum für Naturkunde, Berlin reveals a previously unobserved baby skeleton near the adult. This small skeleton can tell us about how these creatures grew; the lower segment of the hind limb is already proportionally larger than the upper hind limb and the lower and upper forelimbs. This may be because young mesosaurs needed to swim right from birth.
Bryan M. Gee and Robert R. Reisz
Foss. Rec., 21, 79–91, https://doi.org/10.5194/fr-21-79-2018, https://doi.org/10.5194/fr-21-79-2018, 2018
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This paper presents a description of an extremely well-preserved and articulated skeleton of a temnospondyl amphibian from the early Permian of Oklahoma. Postcranial material is not often well-preserved or well-described in the literature for this group, known as dissorophids, and acquiring new data on this region of the skeleton is important for understanding the evolution of the group. It also represents the first documentation at this site of a genus that was otherwise known only from Texas.
Benjamin Ramassamy, Olivier Lambert, Alberto Collareta, Mario Urbina, and Giovanni Bianucci
Foss. Rec., 21, 11–32, https://doi.org/10.5194/fr-21-11-2018, https://doi.org/10.5194/fr-21-11-2018, 2018
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Extant beaked whales perform deep dives to forage for squids. We studied the morphology of the fossil ziphiid Messapicetus gregarius to evaluate its ability to perform such dives. Our analysis suggests an enlargement of the pterygoid sinus system in deep divers. In M. gregarius, the pterygoid sinus is enlarged, but other lines of evidence indicate that the coastal environment also represented an important part of its home range.
Tanja Wintrich, Martin Scaal, and P. Martin Sander
Foss. Rec., 20, 279–290, https://doi.org/10.5194/fr-20-279-2017, https://doi.org/10.5194/fr-20-279-2017, 2017
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Plesiosaurians retain a very long neck but greatly reduce neck flexibility, and the cervicals have large, paired, and highly symmetrical foramina on the ventral side of the centrum, traditionally termed
subcentral foramina, and on the floor of the neural canal. We found that these dorsal and the ventral foramina are connected by a canal. The foramen are not for nutrient transfer; they are the osteological correlates of a highly paedomorphic vascular system in the neck of plesiosaurs.
Volker Lohrmann and Michael S. Engel
Foss. Rec., 20, 239–244, https://doi.org/10.5194/fr-20-239-2017, https://doi.org/10.5194/fr-20-239-2017, 2017
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Amber functions as a window into the past, capable of capturing behaviors
frozenfor millions of years. Here, we report on the exceptionally rare discovery of a dinosaur-age larva of a stinging wasp, feeding on its cricket host, in mid-Cretaceous amber from Myanmar. It reveals a considerable constancy in the biology of this particular family over the last 100 million years. The excellent preservation of the larva is remarkable and due solely to the fidelity permitted by inclusion in amber.
Danilo Harms and Jason A. Dunlop
Foss. Rec., 20, 215–238, https://doi.org/10.5194/fr-20-215-2017, https://doi.org/10.5194/fr-20-215-2017, 2017
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We review the fossil history of pseudoscorpions (Arachnida: Pseudoscorpiones), one of the oldest terrestrial lineages with
a fossil record that goes back to the Devonian. Pseudoscorpions do not fossilise easily, and records from the Mesozoic and
Cenozoic consist almost exclusively of amber inclusions. Overall, 16 of the 26 recent families have been documented from
fossils. This is a group of evolutionary stasis and we discuss aspects of palaeobiology, preservation, and ecology.
Wolfram Mey, Wilfried Wichard, Patrick Müller, and Bo Wang
Foss. Rec., 20, 129–145, https://doi.org/10.5194/fr-20-129-2017, https://doi.org/10.5194/fr-20-129-2017, 2017
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Based on a total of 14 inclusions from Burmese amber the new, extinct insect order Tarachoptera was established. The new order Tarachoptera is placed in the superorder Amphiesmenoptera. The species of Tarachoptera are tiny insects with a wing span of 2.3–4.5 mm, but are highly specialized according to their aberrant morphology. They lived in the forests of Southeast Asia about 100 million years ago.
Florian Witzmann and Elizabeth Brainerd
Foss. Rec., 20, 105–127, https://doi.org/10.5194/fr-20-105-2017, https://doi.org/10.5194/fr-20-105-2017, 2017
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Based on the estimation of its body mass and metabolic rate, we calculated heat balance, gas exchange, osmoregulation, and digestion of the 290-million-year-old aquatic amphibian Archegosaurus. Our results suggest that its physiology was more fish- than tetrapod-like in many respects. We conducted this study, which is mainly based on theoretical calculations and comparison with extant analogs, to shed light on the physiology and lifestyle of early tetrapods.
Torsten M. Scheyer, Elena V. Syromyatnikova, and Igor G. Danilov
Foss. Rec., 20, 69–85, https://doi.org/10.5194/fr-20-69-2017, https://doi.org/10.5194/fr-20-69-2017, 2017
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We review shell bone microstructures of Adocidae and Nanhsiungchelyidae, two groups of extinct hide-necked turtles related to modern soft-shelled turtles (Trionychidae) and the pig-nosed turtle Carettochelys insculpta (Carettochelyidae). Adocids and nanhsiungchelyids both have a good fossil record in North America and Asia during the Cretaceous and Cenozoic. Our microstructural data supplement previously reported differences in external shell morphology between adocids and nanhsiungchelyids.
Olivier Béthoux, Artémis Llamosi, and Séverine Toussaint
Foss. Rec., 20, 1–7, https://doi.org/10.5194/fr-20-1-2016, https://doi.org/10.5194/fr-20-1-2016, 2016
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Fossil insects preserved as an imprint on laminae have subtle differences in relief which can be difficult to photograph. Recourse to reflectance transformation imaging allowed us to deliver exhaustive and interactive photographic data for Protelytron permianum. Based on these data we were able ascertain that it is an early relative of earwigs. In order to better appreciate its elaborate hind wing folding mechanism, we provide guidelines to build and operate a paper model.
Florian Witzmann, Sven Sachs, and Christian J. Nyhuis
Foss. Rec., 19, 83–100, https://doi.org/10.5194/fr-19-83-2016, https://doi.org/10.5194/fr-19-83-2016, 2016
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In this study, the well-preserved skull of a giant amphibian from the Late Triassic Stuttgart Formation (Schilfsandstein) of Bielefeld-Sieker in NW Germany is described and a new species, Cyclotosaurus buechneri sp. nov., is erected. Cyclotosaurus buechneri represents the only unequivocal evidence of Cyclotosaurus in northern Germany. The amphibian skull was found more than 40 years ago and is well known in Bielefeld and the surrounding areas but has so far never been described.
G. Arratia
Foss. Rec., 19, 31–59, https://doi.org/10.5194/fr-19-31-2016, https://doi.org/10.5194/fr-19-31-2016, 2016
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Complete morphological descriptions of exceptionally beautifully preserved Late Jurassic fishes from Bavaria (e.g., Ettling and Eichsttät), southern Germany, are provided for a new genus and species, which is endemic of Ettling and for a new family, Ascalaboidae. The new family is only known from marine Upper Jurassic localities of Europe and is interpreted as an extinct and primitive group of Teleostei, which is considered the largest group within osteichthyans (bony fishes).
N. B. Fröbisch, A. Brar, and R. R. Reisz
Foss. Rec., 18, 73–80, https://doi.org/10.5194/fr-18-73-2015, https://doi.org/10.5194/fr-18-73-2015, 2015
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Here we describe a large specimen of the Paleozoic dissorophid amphibian Cacops woehri from the Fort Sill locality in Oklahoma, which was previously only known from a partial juvenile skull. The new data show that, in contrast to the closely related species C. morrisi, C. woehri underwent only subtle changes in skull morphology in late ontogeny, which indicates different functional demands possibly associated with a different ecology.
B. J. Burger and L. Tackett II
Foss. Rec., 17, 69–74, https://doi.org/10.5194/fr-17-69-2014, https://doi.org/10.5194/fr-17-69-2014, 2014
K. Kienapfel, S. Läbe, and H. Preuschoft
Foss. Rec., 17, 59–67, https://doi.org/10.5194/fr-17-59-2014, https://doi.org/10.5194/fr-17-59-2014, 2014
Cited articles
Aubrechtová, M.: A revision of the Ordovician cephalopod Bactrites sandbergeri Barrande:
Systematic position and palaeobiogeography of Bactroceras, Geobios, 48, 193–211,
https://doi.org/10.1016/j.geobios.2015.03.002, 2015.
Aubrechtová, M. and Turek V.: Lituitid cephalopods from the Middle
Ordovician of Bohemia and their paleobiogeographic affinities, Bull.
Geosci., 93, 401–417, https://doi.org/10.3140/bull.geosci.1707, 2018.
Babcock, L. E.: Trilobites in Paleozoic predator-prey systems, and their
role in reorganization of Early Paleozoic ecosystems, in: Predator-prey
interactions in the fossil record, edited by: Kelley, P. H., Kowalewski, M.,
and Hansen, T. A., Kluwer Academic Publisher, New York, 55–92, 2003.
Bambach, R. K.: Energetics in the global marine fauna: A connection between
terrestrial diversification and change in the marine biosphere, Geobios-Lyon, 32,
131–144, https://doi.org/10.1016/S0016-6995(99)80025-4, 1999.
Barrande, J.: Note sur quelques nouveaux fossils de'couverts aux environs de la ville de Rokitzan dans le bassin silurien du
centre de la Bohême, Bull. Soc. Géol. Fr. Ser. 2, 13, 32–538,
1856.
Barrande, J.: Systême silurien du centre de la Bohême,
Ière Partie: Recherches paléontologiques. Vol. III. Classe des
Mollusques, Ordre des Ptéropodes, Chez l'auteur et éditeur, Prague,
Paris, 1–179 pp. + 16 pls, 1867.
Barrande, J.: Système Silurien du centre de la Bohême, Vol.
I., Chez l'auteur et éditeur, Prague, Paris, I., 1–647, 1872.
Bertling, M.: Arachnostega n. ichnog. – burrowing traces in internal moulds of boring
bivalves (late Jurassic, Northern Germany), Paläont. Z., 66, 177–185,
1982.
Boucot, A. J.: Evolutionary paleobiology of behavior and coevolution, Elsevier, London, 725 pp., 1990.
Boucot, A. J. and Poinar Jr., G. O.: Fossil behavior compendium, CRC Press, Boca Raton, 363 pp., 2010.
Brett, C. E.: Entombment of a trilobite within a closed brachiopod
shell, J. Paleontol., 51, 1041–1045, 1977.
Brett, C. E.: Durophagous predation in Paleozoic marine benthic assemblages,
in: Predator-prey interactions in the fossil record, edited by: Kelley, P.
H., Kowalewski, M., and Hansen, T. A., Kluwer Academic Publisher, 401–432,
2003.
Brett, C. E. and Walker, S. E.: Predators and predation in Paleozoic marine
environments, in The Fossil Record of Predation, edited by: Kowalewski, M.
and Kelley, P. H., Palaeontological Society Special Papers, 8, 93–118, Cambridge University Press, 2002.
Brett, C. E., Zambito, J. J., Hunda, B. R., and Schindler, E.: Mid-Paleozoic
trilobite Lagerstätten: models of diagenetically enhanced obrution
deposits, Palaios, 27, 326–345, https://doi.org/10.2110/palo.2011.p11-040r,
2012.
Bruthansová, J.: The trilobite Family Illaenidae Hawle et Corda, 1847
from the Ordovician of the Prague Basin (Czech Republic), T. Roy. Soc.
Edin-Earth., 93, 167–190, 2003.
Bruthansová, J.: Exuviation of selected Bohemian Ordovician trilobites,
Spec. Pap. Palaeontol., 70, 293–308, 2004.
Bruthansová, J. and Budil, P.: Exuviation of the genus Placoparia Hawle et Corda,
1847 (Trilobita, Czech Republic, Prague Basin, Ordovician), INSUGEO, Ser.
Correl. Geol., 17, 271–274, 2003.
Bruthansová, J., Fatka, O., Budil, P., and Král, J.: 200 years of
trilobite research in the Czech Republic, in: Fabulous Fossils – 300 Years
of Worldwide Research on Trilobites, edited by: Mikulic, M. G., Landing, E.,
and Kluessendorf, J., N. Y. State Mus. Bull., 507, 51–80, 2007.
Budil, P. and Fatka, O.: Frontal Auxiliary Impressions in Ordovician
trilobite genus Dalmanitina Reed, 1905 (Barrandian area, Czech Republic), Bull. Geosci., in press, 2021.
Budil, P., Kraft, P., Kraft, J., and
Fatka, O.: Faunal associations of the Šárka Formation
(Middle Ordovician, Darriwilian, Prague Basin, Czech Republic), Acta
Palaeontol. Sinica Suppl., 46, 64–70, 2007.
Budil, P., Fatka, O., Zwanzig, M., and Rak, Š.: Two
unique Middle Ordovician trilobites from the Prague Basin (Czech Republic),
J. Natl. Mus. Nat. Hist., 179, 95–104, 2010.
Chatterton, B. D. E., Collins, D. H., and
Ludvigsen, R.: Cryptic behaviour in trilobites: Cambrian and
Silurian examples from Canada, and other related occurrences, Spec. Pap.
Palaeontol., 70, 157–173, 2003.
Chatterton, B. D. E. and Fortey, R. A.: Linear cluster sof articulated
trilobites from Lower Ordovician (Arenig) strata at Bini Tinzoulin, North
Zagora, Southern Morocco, in: Advances in trilobite research, edited
by: Rábano, I., Gozalo, R., and García-Bellido, D., Cuadernos del
Museo Geominero, 9: Instituto Geológico y Minero de España, Madrid,
73–77, 2008.
Chlupáč, I.: Neptunian dykes in the Koněprusy Devonian.
Geological and palaeontological observations, Věst. Čes. Geol.
Úst., 71, 193–208, 1996.
Cocks, L. R. M. and Fortey, R. A.: Lower Palaeozoic facies and fauna around
Gondwana, in Gondwana and Tethys, edited by: Audley-Charles, M. G. and
Hallam, A., Geol. Soc. Lond. Spec. Publ., 37, 183–200,
1988.
Davis, R. A., Fraaye, R. H. B., and Holland, C. H.: Trilobites within
nautiloid cephalopods, Lethaia, 34, 37–45, 2001.
Fatka, O. and Budil, P.: Sheltered gregarious behavior of Middle Ordovician
harpetid trilobites, Palaios, 29, 495–500,
https://doi.org/10.2110/palo.2013.031, 2014.
Fatka, O. and Kozák, V.: A new type of entombment of Peronopsis (Agnostida) in a
hyolithid conch, Carnets Geol., 14, 191–198, 2014.
Fatka, O. and Szabad, M.: Agnostids entombed under exoskeletons of
paradoxidid trilobites, N. Jb. Geol. Paläont. Abh., 259, 207–215,
https://doi.org/10.1127/0077-7749/2011/0115, 2011.
Fatka, O., Vokáč, V., Moravec, J., and Valent, M.: Agnostid entombed
in a hyolith conch, Mem. Assoc. Austral. Palaeontol., 37, 481–489, 2009.
Fatka, O., Mikuláš, R., Szabad, M., Micka, V., and Valent, M.: Arachnostega Bertling, 1992 in Drumian (Cambrian) sediments of the Barrandian area (Czech Republic), Acta Geol. Polon., 61, 367–381, 2011.
Fatka, O., Budil, P., and David, M.: Digestive structures in Ordovician
trilobites Colpocoryphe and Flexicalymene from the Barrandian area of Czech Republic, Est. J. Earth
Sci., 64, 255–266, https://doi.org/10.3176/earth.2015.32, 2015.
Fatka, O., Budil, P., and Mikuláš, R.: Healed injury in a
nektobenthic trilobite: “Octopus-like” predatory style in Middle
Ordovican?, Geol. Croat., in press, 2021a.
Fatka, O., Budil, P., and Zicha, O.: Exoskeletal and eye repair in
Dalmanitina socialis (Trilobita): Example of blastemal regeneration in Ordovician?, Int. J.
Paleopathol., in press, 2021b.
Fernández-Álvarez, F. Á., Machordom, A.,
García-Jiménez, R., Salinas-Zavala, C. A., and Villanueva, R.:
Predatory flying squids are detritivores during their early planktonic life,
Sci. Rep.-UK, 8, 3440, https://doi.org/10.1038/s41598-018-21501-y, 2018.
Flick, U. and Flick, H.: Unterkarbon-Trilobiten in Wohnkammern von
Goniatiten – Fossilfalle oder Häutungsplatz?, Der Aufschluss, 60,
245–250, 2009.
Foerse, A. F.: Preliminary notes on Cincinnatian and Lexington fossils of Ohio, Indiana, Kentucky and Tennessee, Bulletin of the Scientific Laboratories of Denison University, 16, 15–87, 1910.
Fortey, R. A.: Early Ordovician trilobite communities, Fossils and Strata, 4,
339–360, 1975.
Fortey, R. A.: Pelagic trilobites as an example of deducing the life habits
of extinct arthropods, T. Roy. Soc. Edin-Earth., 76, 219–230, 1985.
Fortey, R. A. and Owens, R. M.: The Arenig Series in South Wales, Bull.
Brit. Mus. Nat. Hist. Geol., 41, 69–307, 1987.
Fortey, R. A. and Owens, R. M.: Feeding habits in trilobites, Palaeontology,
42, 429–465, 1999.
Gutiérrez-Marco, J.-C., Sá, A. A., García-Bellido, D. C.,
Rábano, I., and Valério, M.: Giant trilobites and trilobite clusters
from the Ordovician of Portugal, Geology, 37, 443–446, https://doi.org/10.1130/G25513A.1, 2009.
Gutiérrez-Marco, J.-C., Sá, A. A., García-Bellido, D. C., and
Rábano, I.: The Bohemo-Iberian regional chronostratigraphical scale for
the Ordovician System and palaeontological correlations within
South Gondwana, Lethaia, 50, 258–295, https://doi.org/10.1111/let.12197,
2017.
Haldeman, S. S.: Supplement to number one of “A monograph of the Limniades, or freshwater univalve shells of North
America”, containing descriptions of apparently new animals in different classes, and the names and characters of the subgenera in Paludina and Anculosa, J. Dobson, Philadelphia, 3 pp., 1840.
Hammann, W.: Die Placopariinae (Trilobita, Cheirurina, Ordovizium),
Senckenberg, Lethaea, 52, 53–75, 1971.
Havlíček, V.: Ordovician of Bohemia: development of the Prague
Basin and its benthic communities, Sbor. Geol. Věd, Geol., 37, 103–136,
1982.
Havlíček, V.: Ordovician, in: Palaeozoic of the Barrandian, edited
by: Chlupáč, I., Havlíček, V., Kříž, J.,
Kukal, Z., and Štorch, P., Czech Geological Survey, Prague, 149–164,
1998.
Havlíček, V. and Vaněk, J.: The biostratigraphy of the
Ordovician of Bohemia, Sbor. Geol. Věd, Paleontol., 8, 7–69, 1966.
Hawle, I. and Corda, A. C. J.: Prodrom einer Monographie der böhmischen
Trilobiten, Abh. Königl. Böhm. Ges. Wiss., 5, 119–292, 1847.
Hicks, H.: On the succession of the Ancient Rocks in the vicinity of St.
David's, Q. J. Geol. Soc. Lond., 31, 167–95, 1875.
Horný, R. J.: Shell breakage and repair in explanate bellerophontoidean
gastropods from the Middle Ordovician of Bohemia, Věst. Čes. Geol.
Úst., 72, 157–168, 1997a.
Horný, R. J.: Shell breakage and repair in Sinuitopsis neglecta (Mollusca, Tergomya) from
the Middle Ordovician of Bohemia, Čas. Nár. Muz., Ř.
Přírodověd., 166, 137–142, 1997b.
Hsieh, A. and Plotnick, R. E.: The representation of animal behaviour in the
fossil record, Anim Behav., 169, 65–80, 2020.
Kelly, P. H., Kowalewski, M., and Hansen, T. A. (Eds.): Predator-prey
interactions in the fossil record: Topics in Geobiology Series 20, Kluwer
Academic Publisher, New York, 1–464, 2003.
Kettner, R. and Kodym, O.: New stratigraphy of the Barrandian, Čas. Mus.
Král. Českého, 93, 47–55, 1919 (in Czech).
Klompmaker, A. A. and Fraaije, R. H. B.: Animal behavior frozen in time:
gregarious behavior of Early Jurassic Lobsters within an Ammonoid body
chamber, PLoS ONE, 7, e31893, https://doi.org/10.1371/journal.pone.0031893,
2012.
Klug, C., Frey, L., Pohle, A., De Baets, K., and Korn, D.: Palaeozoic
evolution of animal mouthparts, Bull. Geosci., 92, 439–442,
https://doi.org/10.3140/bull.geosci.1648, 2017
Kraft, J.: Type specimens of fossils in the West-Bohemian Museum in
Plzeň and in the Regional Museum of Dr. B. Horák in Rokycany. Part I
– Trilobita, Folia musei rerum naturalium Bohemiae occidentalis, Geologica,
1, 1–11, 1972.
Kraft, J. and Kraft, P.: Graptolite biozones of the Bohemian Lower and
Middle Ordovician and their historical development, J. Czech Geol. Soc., 44,
53–62, 1999.
Kraft, P., Kraft, J., and Prokop, R. J.: A possible hydroid from the Lower
and Middle Ordovician of Bohemia, Alcheringa, 25, 143–154,
https://doi.org/10.1080/03115510108619101, 2001.
Kraft, P., Bruthansová, J., and Mikuláš, R.: Feeding traces
related to shells from the Prague Basin, Czech Republic (Tremadocian to
early Darriwilian, Ordovician), Palaeogeography, Palaeoclimatology,
Palaeoecology, 537, 109399, 1–21,
https://doi.org/10.1016/j.palaeo.2019.109399, 2020.
Kröger, B.: Size matters – Analysis of shell repair scars in endocerid cephalopods, Foss. Rec., 14, 109–118, https://doi.org/10.1002/mmng.201100001, 2011.
Kukal, Z.: Petrographical investigation of the Ordovician Šárka beds
in the Barrandian area, Sbor. Ústřed. Úst. Geol., Odd. Geol.,
27, 175–214, 1962 (in Czech, English summary).
Manda, Š.: Trocholites Conrad, 1838 (Nautiloidea, Tarphycerida) in the Middle
Ordovician of the Prague Basin and its palaeobiogeographical significance,
Bull. Geosci., 83, 327–334,
https://doi.org/10.3140/bull.geosci.2008.03.327, 2008.
Marek, L.: The trilobite family Cyclopygidae Raymond in the Ordovician of
Bohemia, Rozpr. Ústřed. Úst. Geol., 28, 1–84, 1961.
Mergl, M., Fatka, O., and Budil, P.: Lower and early Middle Ordovician
trilobite associations of the Prague Basin (Perunica, Czech Republic), Acta
Palaeontol. Sinica Suppl., 46, 320–327, 2007.
Mergl, M., Fatka, O., and Budil, P.: Lower and Middle Ordovician
trilobite associations of Perunica: from shoreface endemicicty to offshore
uniformity (Prague Basin, Czech Republic), in: Advances in trilobite research,
edited by: Rábano, I., Gozalo, R., and García-Bellido, D.,
Cuadernos del Museo Geominero, 9: Instituto Geológico y Minero de
España, Madrid, 275–282, 2008.
Mikuláš, R. and Slavíčková, J.: Trilobites and minute
ovoid pellets in a burrow (Ordovician, Llanvirnian, Czech Republic), Ichnos,
8, 243–249, https://doi.org/10.1080/10420940109380191, 2001.
Mikulic, D. G.: Sheltered molting by trilobites, Geol. Soc. Amer., Boulder. Abstracts with Programs, 26, 55, 1994.
Nixon, M.: The feeding mechanisms and diets of cephalopods – Living and
fossil, in: Cephalopods Present and Past, edited by: Wiedmann J. and
Kullmann, J., 2nd International Cephalopod Symposium, E. Schweizerbart'sche
Verlagsbuchhandlung, Stuttgart, Germany, 642–652, 1988.
Novák, O. P. in Perner, J. (Eds.): Trilobiti D-d1γ z okolí Pražského [Trilobites of D-d1γ from the surroundings of Prague], Palaeontologia Bohemiae, 9, 1–55, 1918 (in Czech).
Prantl, F. and Přibyl, A.: On the Bohemian representatives of the family
Harpetidae (Hawle & Corda), Rozpr. Ústřed. Úst. Geol., 18,
1–170, 1954 (in Czech, Russian and English summary).
Owens, R. M. and Servais, T.: The Ordovician of the Condroz Inlier, Belgium:
Trilobites from the southeastern margin of Avalonia, Palaeogeogr. Palaeoecl.,
245, 272–294, 2007.
Peder, A. E. H. and Feist, R.: Lower Devonian (Emsian) rugosa of the Izarne
Formation, Montagne Noire, France, J. Paleontol., 72, 967–991, 1998.
Přibyl, A. and Vaněk, J.: Einige Trilobiten aus dem böhmischen
Ordovizium, Věst. Ústřed. Úst. Geol., 43, 191–197, 1968.
Radwański, A., Kin, A., and Radwańska, U.: Queues of blind phacopid
trilobites Trimerocephalus: A case of frozen behaviour of Early Famennian age from the Holy
Cross Mountains, Central Poland, Acta Geol. Pol., 59, 459–481, 2009.
Rakociński, M.: Phenomenon of “sheltered preservation” – an overview
and fossil examples from the Upper Devonian of Poland, Przeglad Geol., 57,
584–590, 2009.
Raymond, P. E.: Note on the names Amphion, Harpina and Platymetopus, Am. J. Sci., 19, 377–378, 1905.
Salter, J. W.: On the fossils of the Lingula-Flags or “Zone Primordiale”, Quarterly Journal of the Geological Society of London, 15, 551–555, 1859.
Schoenemann, B., Clarkson, E. N. K., and Hoyberget, M.: Traces of an ancient
immune system – how an injured arthropod survived 465 million years ago,
Sci. Rep.-UK, 7, 40330, https://doi.org/10.1038/srep40330, 2017.
Šnajdr, M.: Note on the regenerative ability of injured trilobites,
Věst. Ústřed. Úst. Geol., 54, 171–173, 1979.
Struve, W.: Beitrage zur Kenntnis der Phacopacea (Trilobita) 1: Die Zeliszkellinae, Senckenbergiana
lethaea, 39, 165–219, 1958.
Suzuki, Y. and Bergström, J.: Trilobite taphonomy and ecology in Upper
Ordovician carbonate buildups in Dalarna, Sweden, Lethaia, 32, 159–172,
https://doi.org/10.1080/11035890903452670, 1999.
Valent, M., Fatka, O., Micka, V., and Šinágl, M.: Hyoliths with
entombed trilobites – cryptic behavior of trilobites?, in: Advances in
trilobite research, edited by: Rábano, I., Gozalo, R., and
García-Bellido, D., Cuadernos del Museo Geominero, 9: Instituto
Geológico y Minero de España, Madrid, 411–413, 2008.
Vinn, O., Wilson, M. A., Zatoń, M., and Toom, U.: The trace fossil
Arachnostega in the Ordovician of Estonia (Baltica), Palaeontol. Electron., 17, 1–9,
2014.
Vokáč, V., Hartl, F., David, M., Pavlovič, M., Doubrava, M.,
Kozák, V., and Grigar, L.: Notable findings of trilobites from the
Ordovician (Dapingian–Sandbian) of the Prague Basin (Barrandian area, Czech
Republic), Erica, 22, 141–157, 2015 (in Czech, English summary).
Vokáč, V., Hartl, F., Pavlovič, M., Beneš, P., Zicha, O.,
Grigar, L., Tichávek, F., and Henkl, L.: Remarkable new findings of
Ordovician (Darriwillian – Sandbian) trilobites in the southwest part of
the Prague Basin (Rokycany region, Czech Republic), Erica, 26, 67–93, 2019
(in Czech, English summary).
Wani, R.: How to recognize in situ fossil cephalopods: evidence from
experiments with modern Nautilus, Lethaia, 40, 305–311, 2007.
Short summary
Sixteen samples contain one to six Ordovician trilobites preserved in the protected space inside of empty cephalopod conchs or under diverse parts of exoskeletons of much larger disarticulated trilobites. Analyses of the level of articulation of trilobites in combination with their orientation, disposition, and placement make it possible to conclude that the hidden trilobites deliberately entered shells of dead cephalopods for food, refuge, ecdysis, reproduction or lodging.
Sixteen samples contain one to six Ordovician trilobites preserved in the protected space inside...