Articles | Volume 24, issue 2
Research article
05 Jan 2022
Research article |  | 05 Jan 2022

Ingensalinae subfam. nov. (Hemiptera: Fulgoromorpha: Fulgoroidea: Inoderbidae), a new planthopper subfamily from mid-Cretaceous Kachin amber from Myanmar

Cihang Luo, Zhishun Song, Xiaojing Liu, Tian Jiang, Edmund A. Jarzembowski, and Jacek Szwedo

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Cited articles

Bartlett, C. R., Deitz, L. L., Dmitriev, D. A., Sanborn, A. F., Soulier-Perkins, A., and Wallace, M. S.: The diversity of the true hoppers (Hemiptera: Auchenorrhyncha), in: Insect biodiversity, Science and society, 2, edited by: Foottit, R. G. and Adler, P. H., Wiley-Blackwell, Oxford, 501–590,, 2018. 
Bourgoin, T.: FLOW (Fulgoromorpha Lists on the Web): a world knowledge base dedicated to Fulgoromorpha, Version 8, updated: 19 December 2021, available at:, last access: 20 December 2021. 
Bourgoin, T. and Campbell, B. C.: Inferring a phylogeny for Hemiptera: falling into the “autapomorphic trap”, in: zugleich Kataloge des OÖ, Landesmuseums, Neue Folge Nr. 176, Denisia 04, 67–82, 2002. 
Bourgoin, T. and Szwedo, J.: The “cixiid-like” fossil planthopper families, Bull. Insectol., 61, 107–108, 2008. 
Bourgoin, T., Wang, R.-R., Asche, M., Hoch, H., Soulier-Perkins, A., Stroiński, A., Yap, S., and Szwedo, J.: From micropterism to hyperpterism: recognition strategy and standardized homology-driven terminology of the forewing venation patterns in planthoppers (Hemiptera: Fulgoromorpha), Zoomorphology, 134, 63–77,, 2015. 
Short summary
Ingensala xiai gen. et sp. nov. is described based on a specimen from mid-Cretaceous Burmese amber. The new genus can be definitely attributed to Inoderbidae mainly based on its head structure, pronotum, and mesonotum without median and lateral carinae, but it differs from the type genus to a large extent and two new subfamilies are established for these two genera respectively. The similarities between Ingensala, Qiyangiricaniidae, Weiwoboidae, and Tropiduchidae are discussed.