Articles | Volume 20, issue 1
Research article
21 Feb 2017
Research article |  | 21 Feb 2017

A re-examination of the enigmatic Russian tetrapod Phreatophasma aenigmaticum and its evolutionary implications

Neil Brocklehurst and Jörg Fröbisch

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Cited articles

Aitchison, J.: The Statistical Analysis of Compositional Data, Chapman & Hall, London, 1986.
Benson, R. B. J.: The global interrelationships of basal synapsids: cranial and postcranial morphological partitions suggest different topologies, J. Syst. Palaeontol., 10, 601–624, 2012.
Benton, M. J.: No gap in the Middle Permian record of terrestrial vertebrates, Geology, 40, 339–342, 2012.
Berman, D. S., Henrici, A. C., Kissel, R. A., Sumida, S. S., and Martens, T.: A new diadectid (Diadectomorpha), Orobates pabsti, from the Early Permian of central Germany, Bulletin of the Carnegie Museum of Natural History, 35, 1–36, 2004.
Botha-Brink, J. and Modesto, S. P.: Anatomy and relationships of the Middle Permian varanopid Heleosaurus scholtzi based on a social aggregation from the Karoo Basin of South Africa, J. Vertebr. Paleontol., 29, 389–400, 2009.
Short summary
Phreatophasma is a mysterious species from the middle Permian of Russia. Since its description in 1954, it has been assigned to many different groups: therapsids (the most diverse of middle Permian terrestrial vertebrates), caseids (one of the earliest herbivore lineages), and seymouriamorphs (a Permian amphibian clade). Using recent knowledge of morphology and state-of-the art analyses, we confirm that Phreatophasma is one of the youngest members of Caseidae, the second member to enter Russia.